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Botany of the wheat plant. A wide diversity of topics, ranging from the fine structure of. Modern agronomic methods, however, are often. Some researchers will require more detailed.

MATURE PLANTThe . All the. structures, such as leaves, tillers and ears, have attained their full size. Plant. The plant is made up of a root and shoot system. Two types of. roots are found, the seminal roots and the nodal roots (adventitious or crown. The shoot is made up of a. Figure 2. 1). There are from 6 to 1.

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From four to seven of the most distal units have. The portion of the shoot with elongated internodes is. In the proximal or basal units, the internode. A leaf is inserted at. The shoot is terminated by an ear or spike bearing about 2. In the ear, the phytomer is made up of the spikelet (the axillary. Tillers. Tillers, which have the same basic structure as the main.

At anthesis, only some of the. Others die and may be.

Leaves. Each leaf comprises the sheath, wrapping around the subtending. At the junction of the sheath and lamina, there is a. The base of the leaves on the culm is thickened to form a hard knot or. Internodes. The elongated distal internodes increase in length from the.

Figure 2. 1). FIGURE 2. Diagram of a 'mature' plant, alternative numbering system for leaves and internodes. Nomenclature. Different systems for identifying leaves and internodes have. As with. the total number of leaves, the number of nodes on the shoot are often not known. Some systems use the uppermost leaf (the flag leaf) or the.

Other systems are used in development studies where the position and. Figure 2. 1). THE SEED, GERMINATION AND SEEDLING EMERGENCEThe seed, grain or kernel of wheat (more pedantically, the. The dorsal side (with respect to the. Figure 2. 2). The embryo or germ is situated at the point of attachment of the. The Fem Bot watch with english subtitles in 2160 21:9.

The embryo is made. Figure. 2. 3). The scutellum is the region that secretes some of the enzymes involved in. Surrounding the endosperm is a metabolically active layer of cells or. In. the plumule region of the embryo, the coleoptile, about four leaf primordia and. FIGURE 2. 2. Wheat grain, showing different aspects and cross section to illustrate the depth. The coleoptile is well developed in the embryo, forming a.

At. germination, it bursts through the pericarp and testa and grows through the soil. It is white in colour except. The end of the coleoptile is bullet- shaped and closed except for a. Two vascular bundles are found laterally placed with reference.

One or two lines of stomata extending from. They tend to be more numerous towards the tip. Stomata also. occur in the inner epidermis though they are less frequent. The guard cells of. The other cells of the outer epidermis have thickened walls. The inner epidermis has an unthickened wall, which is not wrinkled.

The radicle emerges first and then the plumule. As growth continues. The coleoptile emerges. The coleoptile increases in length until it emerges.

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If the seed is sown. In wheat, unlike oats and. In the case of very deeply sown seeds (more than 1. ROOTSThe wheat plant has two types of roots, the seminal (seed). About six root primordia are present in the embryo. At. germination, the primary root bursts through the coleorhiza, followed by the. These form the seminal root.

Nodal roots are associated with tiller development and are usually. Compared with the.

Nodal. roots occur on the lower three to seven nodes (depending on environmental. The uppermost node on which. At. maturity, the root system extends to between 1 and 2 m deep or more depending on. Most roots occur in the top 3. FIGURE 2. 4. A deeply sown seeding (left) with coleoptile removed, showing the development.

Source: Kirby and Appleyard. Courtesy of Arable Unit RASE))Nomenclature.

E.j. Bellocq

While the general classification of seminal and nodal roots is. Klepper et al., 1. At. each node, a root arises from one of four quadrants, which may be designated X.

Y, A and. B. Using this system, any root can be identified by the node at which it arises. Thus, root 2. X is the root arising at node (leaf) 2 in the leaf midrib.

Klepper et al., 1. Anatomy. The young seminal root has a root cap behind which the root is. A transverse section of the root in the root hair. The walls of the cells at the cortex- stele interface (the. The stele has alternate bands of.

Esau. 1. 95. 3). Further back from the root hair zone, lateral branch roots. They grow through the. In the older regions of the root, the cortex dies leaving only the stele. In the deeper. regions of the soil, the anatomy of the nodal roots is similar to that of a. Where the roots emerge from the node, near the soil surface, the. The. region of thickening extends for about 4. This anchors the plant firmly in.

When lodging occurs through the. Ennos, 1. 99. 1). SHOOT APEX DEVELOPMENTThe apex is already well formed in the embryo, and from. Bonnet, 1. 96. 6; Gardner et al., 1. Kirby and. Appleyard, 1.

Williams, 1. 97. 5). The change in the form of the apex results. Kirby, 1. 97. 4). At first the apex is conical in shape and initiates leaves. Figure 2. 5a). As development proceeds the apex becomes more cylindrical in.

Figure. 2. 5b). The next recognizable stage is the double ridge stage (Figure 2. Each. of the primordia initiated on what will eventually become the ear has two parts.

The lower, smaller ridge is a leaf primordium, the further. The upper, larger ridge eventually. The double ridge stage is sometimes. From the double ridge stage onwards, attention is focused on. Figure. 2. 5d- f). Eventually, when about 2.

This occurs when the last initiated primordia, instead of. Figure. 2. 5g). The terminal spikelet stage is regarded as a key stage in wheat. At the terminal spikelet stage, the shoot apex is a fully formed.

Leaf. primordia arise from the superficial layer of cells (dermatogen, tunica), while. Figure 2. 6). LEAVESThe leaf. The leaf is divided at the ligule into a cylindrical sheath. The sheath is tubular at the base, but nearer to. The lamina has a fairly.

It. divides the blade into two subequal parts, each of which has a number of. Each vein marks the position of a vascular. The abaxial surface is more or less. The midrib extends down into the sheath for a short distance as a. The leaf blade naturally assumes a twist, and just below the. This constriction is produced by the constraint upon growth. The ligule is a thin colourless flap of tissue about 1 to 2 mm in.

Associated with the ligule are the auricles, two small earlike. FIGURE 2. 5. Successive stages of shoot apex development from a vegetative apex (a) to. Source: Adapted from Kirby and Appleyard, 1. Courtesy of Arable Unit RASE)FIGURE 2. Longitudinal section of shoot apex at the double ridge stage. At the base of the leaf sheath of the culm leaves, there is a. It is considerably.

The. node of the stem is below the joint and its position is marked by a slight. The joint has an important function, lifting the ear. Leaf shape and size change with leaf position. The lowermost. leaf on the main shoot has parallel sides to within 1 cm or so of the tip so. The leaves above the first have. The last leaf produced upon the culm, the flag leaf.

In spring wheat, lamina length increases with increasing leaf. Lamina width increases with leaf position so that the flag leaf is. Sheath length also increases with leaf position, markedly so.

Winter wheat shows comparable size changes, but associated. The. ad- and abaxial epidermis of the mature leaf enclose the mesophyll, which is. Esau, 1. 95. 3). The vascular tissue. The vascular tissue lies beneath the. Epidermis. The adaxial epidermis is a complex tissue with several cell. Esau, 1. 95. 3). The bulliform (bubble- shaped) cells are the largest cells. When seen in optical.

Flanking the bulliform cells are long. There is usually a single row of stomata.

Each stoma is made. The stoma length varies from 4. The frequency of stomata. There are more on the adaxial surface and are. On the other flank of the row of.

The long cells are interspersed in a regular manner by. Short, unicellular hairs.

The abaxial epidermis has fewer cell types, mainly the long. Stomata occur in. The epidermis on both surfaces of the leaf has a cuticle with. The form of the wax depends upon the. This occurs as. lobed plates, simple plates, flat ribbons and tubes, the amount and form of the. Mesophyll. The mesophyll cells are of a complex lobed shape, resembling. Esau, 1. 95. 3). When viewed in transverse section, the.

The. cells in the middle layers of the leaf are not so elongated. Viewed in. longitudinal section, the lobed nature of these cells is apparent. In the leaves. at the base of the plant, the degree of lobing is low and the dimensions of the.

With ascending leaf position up the stem, the degree of lobing. The effect of these changes. There is variation in the compactness and arrangement of the.

Some cultivars have a relatively loose arrangement of cells. Prominent sub- stomatal cavities. Vascular tissue. The vascular bundle has the structure typical of a.

C3 plant. The phloem is abaxial to the xylem and in the larger. The. xylem has two large, prominent xylem vessels between which are smaller metaxylem. Adaxial to the metaxylem, there is an area of disrupted. The conducting elements are surrounded by an inner (mestome) sheath.

C3 plants. The cells of the mestome sheath are small and. Those of the outer bundle sheath are. In longitudinal section, the. The walls of the. The complex fine structure. O’Brien and Zee, 1. The small veins that interconnect the main longitudinal veins.

They pass through the mestome and parenchyma sheaths and connect directly. They do not have bundle. Vascular system. The major bundles run parallel with each other the whole. The small transverse veins, which constitute about 7 percent. Figure 2. 7). Towards the tip of. Y- shape, the forks of.

Y comprising small transverse veins that link to the longitudinal veins at. At the pointed tip of the leaf, the veins converge and connect with. The distance between the longitudinal veins varies from about 0. Black- man, 1. 97.

Leaf sheath base.

Bellocq - Wikipedia. E. J. Bellocq. Bellocq in 1. Born. John Ernest Joseph Bellocq. New Orleans. Died. New Orleans. Resting place. New Orleans. Nationality.

American. Occupation. Photographer. Years active. John Ernest Joseph Bellocq (1.

American professional photographer who worked in New Orleans during the early 2. Bellocq is remembered for his haunting photographs of the prostitutes of Storyville, New Orleans' legalized red light district. These have inspired novels, poems and films. Bellocq was born into a wealthy white French Creole family in the French Quarter of New Orleans. He became known locally as an amateur photographer before setting himself up as a professional, making his living mostly by taking photographic records of landmarks and of ships and machinery for local companies. However, he also took personal photographs of the hidden side of local life, notably the opium dens in Chinatown and the prostitutes of Storyville.

These were only known to a small number of his acquaintances. In the latter part of his life, he lived alone and acquired a reputation for eccentricity and unfriendliness. According to acquaintances from that period, he showed little interest in anything other than photography. In his early days, he had been something of a dandy. Bellocq photograph circa 1.

Bellocq died in 1. Saint Louis Cemetery #3 in New Orleans. After his death, most of his negatives and prints were destroyed. However, the Storyville negatives were later found.

After many years, they were purchased by a young photographer, Lee Friedlander. In 1. 97. 0, a show of Friedlander's posthumous prints on gold tone printing out paper from Bellocq's 8. A selection of the photographs were also published concurrently in the book, Storyville Portraits.

These photographs were immediately acclaimed for their unique poignancy and beauty. A more extensive collection of Friedlander's prints, entitled Bellocq: Photographs from Storyville, was published with an introduction by Susan Sontag in 1. In recent times, a significant number of prints from Bellocq's own studio have come to light. They are typical professional photographs of the day, such as portraits, copy work for the Louisiana State Museum, and local views, yet few if any Storyville portraits printed by Bellocq's hand exist. A number of early posthumous prints from Bellocq's negatives by photographer Dan Leyrer have also surfaced.

Bellocq Gallery of Photography at Louisiana Tech University is named in his honor. The Storyville photographs. Some are nude, some dressed, others posed as if acting a mysterious narrative.

Many of the negatives were badly damaged, in part deliberately, which encouraged speculation. Many of the faces had been scraped out; whether this was done by Bellocq, his Jesuit priest brother who inherited them after E. J.'s death or someone else is unknown. Bellocq is the most likely candidate, since the damage was done while the emulsion was still wet. In a few photographs the women wore masks.

Some prints made by Bellocq have since surfaced. These are far more conventional than the full- negative prints made by Friedlander. The Storyville photographs not only serve as a record of the prostitutes, but also the interiors of the businesses that housed them.

Bellocq in literature and film. He also appears in Michael Ondaatje's novel Coming Through Slaughter and is a protagonist in Peter Everett's novel Bellocq's Women. These works take many liberties with the facts of Bellocq's life.

He is also a minor character in David Fulmer's novel Chasing the Devil's Tail. The photographs have inspired imaginative literature about the women in them. There are several collections of poems, notably Brooke Bergan's Storyville: A Hidden Mirror and Natasha Trethewey's Bellocq's Ophelia. The 1. 97. 4 book Storyville, New Orleans: Being an Authentic, Illustrated Account of the Notorious Red- Light District by Al Rose gives an overview of the history of prostitution in New Orleans with many photographs by Bellocq.

In 1. 97. 1, . Wright Campbell features a thinly- veiled depiction of Bellocq, a photographer named B. E. Locque. Bellocq appears as a fictional character in David Fulmer's Storyville novels Chasing the Devil's Tail and Rampart Street. He is also a character in Madam: A Novel of New Orleans by Cari Lynn and Kellie Martin.